Running head : Striatum signaling physical effort 1 1 Choosing to make an effort : the role of striatum in signaling physical effort of a chosen

1 Choosing to make an effort: the role of striatum in signaling physical effort of a chosen 1 action 2 3 Kurniawan, I.T., Seymour, B., Talmi, D., Yoshida, W., Chater, N., & Dolan, R. J. 4 5 Cognitive, Perceptual, and Brain Sciences 6 University College London 7 Gower Street, London WC1E 6BT, UK 8 9 Wellcome Trust Centre for Neuroimaging 10 University College London 11 12 Queen Square, London WC1N 3BG, UK 12 13 ESRC Centre for Economic Learning and Social Evolution (ELSE) 14 Drayton House, Gordon Street, London WC1H OAN, UK 15 Department of Economics, University College London 16 Gower Street, London WC1E 6BT, UK 17 18 Corresponding author: Irma T. Kurniawan 19 Cognitive, Perceptual, and Brain Sciences 20 University College London 21 Gower Street, London WC1E 6BT, UK 22 23 Tel: +44 207 679 5379 24 Fax: +44 207 436 4276 25 E-mail: [email protected] 26 27 28 No. of figures : 5 29 No. of tables : 3 30 No. of pages : 33 31 32 33 34 35 36 37 38 39 40 Acknowledgments: The authors would like to thank Eric Featherstone and Alphonso Reid for 41 their technical support during experimental preparation. DT is currently at the School of 42 Psychological Sciences, University of Manchester, UK. 43 44 Grants: This work was supported by a Wellcome Trust Programme Grant to R.J.D. N.C. is 45 supported by a Leverhulme Trust Major Research Fellowship and the ELSE Research Centre at 46 UCL. 47 Articles in PresS. J Neurophysiol (May 12, 2010). doi:10.1152/jn.00027.2010